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Related: Was Darwin Wrong?
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Editor's Note
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The Man Who Wasn't Darwin
Alfred Russel Wallace charted a great dividing line in the living world—and found his own route to the theory of evolution.

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Photograph by Robert Clark; Sophia M. Sachs Butterfly House, Missouri Botanical Garden
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Photograph by Robert Clark; Cleveland Metroparks Zoo
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Photograph by Robert Clark; Natural History Museum, London
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Photograph by Mattias Klum
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Photograph by Robert Clark; Natural History Museum, London
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Photograph by Robert Clark; Natural History Museum, London
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Photograph by Robert Clark; Fort Wayne Children's Zoo
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Photograph by Robert Clark; Fort Wayne Children's Zoo
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Photograph by Robert Clark; Natural History Museum, London
Ornithoptera croesus, show the sort of subtle individual variations that provide the raw material for natural selection. Wallace collected more than a hundred specimens on an island called Batchian.]]>
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Photograph by Robert Clark; Natural History Museum, Tring
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Photograph by Robert Clark; Fort Wayne Children's Zoo
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Photograph by Robert Clark; Saint Louis Zoo
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Photograph by Robert Clark; Santa Barbara Zoo
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Photograph by Robert Clark; Natural History Museum, Tring
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Photograph by Robert Clark; Natural History Museum, Wandsworth
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Photograph by Robert Clark; Sophia M. Sachs Butterfly House, Missouri Botanical Garden
Ornithoptera priamus
The streamlined wings of birdwing butterflies prompted Wallace to speculate on their usefulness and the natural conditions that shaped them. "A short and rounded wing," he wrote, "accompanies a more feeble or more laborious flight," while pointed wings, like those found on fast-flying terns and falcons, give "increased rapidity." At some point, Wallace thought, the island habitat of members of the Papilionidae, or swallowtail family, must have been shared with such an abundance of insect-eaters that the threat of being consumed made "some unusual means of escape a necessity for the large-winged and showy butterflies." ]]>
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Photograph by Robert Clark; Bronx Zoo
Paradisaea minor
Extravagantly feathered birds of paradise captivated Wallace. When he published a two-volume account of his travels through the Malay Archipelago in 1869, he declared himself "the only Englishman who has seen these wonderful birds in their native forests."]]>
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Photograph by Robert Clark; Division of Birds, Smithsonian Institution
Pittagenus (all)
Wallace spotted many species of pittas on his travels. These specimens are all from Sumatra and Borneo, west of Wallace's line; some species' ranges extend far beyond the island where they were collected. Wherever he found the plump tropical birds, Wallace wrote, pittas shared the habit of hopping across the ground in search of insects and seemed "free to adorn themselves with the brightest hues from Nature's laboratory."]]>
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Photograph by Robert Clark; Division of Mammals, Smithsonian Institution
Babyrousa celebensis
Several species of babirusa, each having four curving tusks, long legs, and a largely fruit-based diet, are indigenous to Sulawesi and a few neighboring islands. The babirusa, Wallace wrote, "stands completely isolated, having no resemblance to the pigs of any other part of the world."]]>
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Photograph by Robert Clark; Natural History Museum, London
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